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Explain the principles of cohesion tension theory and its limitations?

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1. Cohesion tension theory is widely accepted theory for translocation of water proposed by Dixon and Joly. 

2. It is based on principles cohesion and adhesion of water molecules along with transpiration pull. 

3. Strong attractive force between water molecules is cohesive force and the strong force of attraction between water molecules and wall of lumen of xylem vessels is called adhesive force. 

4. Owing to these cohesive and adhesive forces a continuous column of water is maintained in xylem elements from root to aerial parts, tips of leaves. 

5. Transpiration pull is developed in the xylem vessel in leaves. 

6. This pull is transmitted downwards and due to suction pressure, water is pulled passively against gravity, i.e. ascent of sap.

Limitations of theory-

1. For activity of transpiration pull, water column should be maintained constantly and continuously. Owing to changes in temperature during day and night, gas bubbles may be formed in the water channel. This will break the continuity. 

2. According to this theory, tracheids are more efficient than vessels because of their tapering end walls which support water column. Vessels are tubular structure with open ends.

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